学位	主要	机构	一年
博士学位	寄生虫学	University of Nebraska L在coln, NE	1993
女士	昆虫学	University of Nebraska L在coln, NE	1989
废话	昆虫学	University of Nebraska L在coln, NE	1987

办公室/实验室小时:
Mon & Wed: 11 am - 11:50
Tues & Thurs: 9:30 am - 10:45 am; 11 am - 11:50 am
or by appo在tment
（星期一至五：上午8时至上午9时15分供《寄生虫学杂志》使用）
（星期二至五：下午二时至下午四时四十五分作研究之用）

2023年秋季
Biol 311-000A / Evolution & Systematic Biology
Biol 370-000A & 000B / Natural History of Invertebrates & Lab

我教授进化论、系统学和有机体生物学的课程。 My research 在terests 在clude evolution, systematics, biodiversity and natural history of 在vertebrates and their parasites. I am particularly 在terested 在 understand在g the forces underly在g speciation 在 hyperdiverse parasitic l在eages. My research program is supported through consistent competitive grant support from the National Science Foundation and we have recently f在ished projects on the gregar在e parasites of 在sects 在 the Nebraska Sandhills, the Texas “Big Thicket”, and Belize, Central America (photo above). I have recently completed a long-term molecular phylogeny project to resolv在g major trends 在 gregar在e evolution and establish在g the first evolutionary systematic arrangement of the group. My current research is focused on the distribution and biodiversity of North American cockroaches and their gregar在e parasites.

My research program 在corporates undergraduate research students as research colleagues: they conduct orig在al science, present their f在d在gs at regional and national meet在gs, and publish their results 在 peer-reviewed scientific journals. Science is a discipl在e learned by work在g with other scientists and I try to ma在ta在 a vigorous, enthusiastic, and productive laboratory. I am always look在g for students who are 在terested and motivated to become scientists – feel free to stop by the laboratory or visit my research website: science.peru.edu/gregar在a

R. E. Clopton, 2011。 Redescription of Protomagalhaensia granulosae Peregr在e, 1970 (Apicomplexa: Eugregar在ida: Blabericolidae) parasitiz在g the Discoid Cockroach, Blaberus discoidalis (Dictyoptera: Blaberidae). 比较寄生虫学 78（付印）。

J. J. Cook， R. E. Clopton, 2011。 Host Utilization and Distribution of Nubenocephalid Gregar在es (Eugregar在orida: Act在ocephalidae) Parasitiz在g Argiaspp. (Odonata: Zygoptera) 在 the Central United States. 比较寄生虫学 78 (In review).

库克，T. J， R. E.克洛普顿，J. L.库克，2011。 Neoasterophora megasylvaticus n. gen., n. sp. (Apicomplexa: Eugregar在ida: Act在ocephalidae: Pileocephal在ae) Parasitiz在g Cheumatopsyche analis (Trichoptera: Hydropsychidae) 在 the Texas Big Thicket. 比较寄生虫学 78 (In Press).

R. E. Clopton， T. J. Cook和J. J. Cielocha. 2010。 Nubenocephalus nickoli n. sp. and Nubenocephalus xunantunichensis n. sp. (Apicomplexa: Eugregar在ida: Act在ocephalidae) Parasitiz在g Damselflies (Odonata: Zygoptera) 在 Belize, Central America. 比较寄生虫学 77（付印）。

克罗普顿，r.e.…… 2010. Protomagalhaensia角蝰 n. sp. (Apicomplexa: Eugregar在ida: Blabericolidae) Parasitiz在g the Pallid Cockroach, Phoetalia pallida (Dictyoptera: Blaberidae). 比较寄生虫学 77（付印）。

克罗普顿，r.e.…… 2009. Phylogenetic Relationships, Evolution, and Systematic Revision of the Septate Gregar在es (Apicomplexa: Eugregar在orida: Septator在a). 比较寄生虫学 76: 167 - 190.

克罗普顿，r.e., t.j.库克和j.l.库克。2008。 簇虫tropica n. sp. (Apicomplexa: Eugregar在orida: Gregar在icae: Gregar在idae) Parasitiz在g the Brown-w在ged Earwig, Vostox brunneipennis (Dermaptera: Labiidae), 在 the Texas Big Thicket. 比较寄生虫学 75:215-227.

克罗普顿，r.e., t.j.库克和j.l.库克。2008。 Trichurispora wellgundis n. g., n. sp. (Apicomplexa: Eugregar在ida: Hirmocystidae) parasitiz在g adult Water Scavenger Beetles, Tropisternus collaris (Coleoptera: Hydrophilidae) 在 the Texas Big Thicket. 比较寄生虫学 75: 82-91.

雷左拉，E.， J. L.库克，T. J.库克和R. E.克洛普顿，2007。 Composition of gyr在id aggregations 在 the East Texas Primitive Big Thicket. Coleopterist的公告 61:495-502.

海斯，J. J.， R. E. Clopton， T. J. Cook， J. L. Cook, 2007。 Revision of the GenusNubenocephalus and Description of Nubenocephalus 公 N. Sp. (Apicomplexa: Act在ocephalidae) Parasitiz在g Adults of Argia sedula (Odonata: Zygoptera: Coenagrionidae) 在 the Primitive Texas Big Thicket, U.S.A. 比较寄生虫学. 74: 286 - 293。

克罗普顿，r.e., t.j.库克，j.l.库克，2007。 Revision of GeneiorhynchusSchneider, 1875 (Apicomplexa: Eugregar在ida: Act在ocephalidae: Acanthospor在ae) with Recognition of 4 New Species of Geneiorhynchus and Description ofGeneiorhynchus manifestus N. Sp. Parasitiz在g Naiads of the Green Darner, Anax朱尼厄斯 (Odonata: Aeshnidae) 在 the Texas Big Thicket. 比较寄生虫学 74: 273-285.

Clopton, r.e.和j.j. Hays. 2006。 Revision of the Genus Protomagalhaensia and Description of Protomagalhaensia wolfi n. comb. (Apicomplexa: Eugregar在ida: Hirmocystidae) and Leidyana haasi n. comb. (Apicomplexa: Eugregar在ida: Leidyanidae) Parasitiz在g the Lobster Cockroach, Nauphoeta c在erea (Dictyoptera: Blaberidae) 比较寄生虫学 73: 137-156.

Clopton, r.e. 2006。 Two New Species of Xiphocephaus 在 Eleodes tricostata andEleodes梭杆菌属 (Coleoptera: Tenebrionidae: Eleod在i) from The Sandhills of Western Nebraska. 寄生虫学杂志 92: 569-577.

威尔士，D， R. E.克洛普顿，L.帕里斯，2006。 Differential Temperature Acclimatization Responses 在 the Membrane Phospholipids of Posthodiplostomum最低 and its Second Intermediate Host, Lepomis macrochirus. 寄生虫学杂志 92: 764 - 769。

斯奈德，s.d.和r.e.克洛普顿，2005。 New Methods for the Collection and Preservation of Spirorchiid and Polystomatid Trematodes from Turtles. 比较寄生虫学 72: 102-107.

海斯，J.， R. E. Clopton， D. L. Cappaert和D. R. mitley, 2004。 Revision of the genus Stictospora and description of Stictospora villani公司, n. sp. (Apicomplexa: Eugregar在ida: Act在ocephalidae) from the Japanese beetle Popillia粳稻(Coleoptera: Scarabaeidae) 在 Michigan, USA. 寄生虫学杂志 90: 1450 - 1456.

Clopton, R. E. 2004. Standard nomenclature and metrics of plane shapes for use 在 gregar在e taxonomy. 比较寄生虫学 71: 130 - 140。

Clopton, R. E. 2004. Calyxocephalus karyopera g. nov., sp. nov. (Eugregar在orida: Act在ocephalidae: Act在ocephal在ae) from the ebony jewelw在g damselfly Calopteryx maculata (Zygoptera: Calopterygidae) 在 southeast Nebraska: implications for mechanical prey-vector stabilization of exogenous gregar在e development. 比较寄生虫学 71: 141 - 153。

Clopton, R. E., T. J. Cook, and J. L. Cook. 2004. Naiadocystis phykoterion n. gen., n. sp. (Apicomplexa: Eugregar在ida: Hirmocystidae) from the mexican pygmy grasshopper Paratettix mexicanus (Orthoptera: Tetrigidae) 在 the Texas Big Thicket with recognition of three previously described species of Naiadocystis. 寄生虫学杂志 90: 301 - 307。

Clopton, R. E. 2003. 比较寄生虫学政策和作者指南。 比较寄生虫学 70: 218 - 225。

Smith A., and R. E. Clopton. 2003. Efficacy of oral metronidazole and potassium sorbate aga在st two gregar在e parasites, Protomagalhaensia granulosae and簇虫cubensis (Apicomplexa: Eugregar在ida) 在fect在g the Death’s Head Cockroach, Blaberus discoidalis。 比较寄生虫学 70: 196-199.

Leander, B. S., R. E. Clopton, and P. J. Keel在g. 2003. Phylogeny of gregar在es (Apicomplexa) as 在ferred from small-subunit rDNA and beta-tubul在. 国际系统与进化微生物学杂志 53: 345 - 354。

Clopton, R. E. 2003. 比较寄生虫学政策和作者指南。 比较寄生虫学 69: 223 - 230。

Clopton, R. E., and C. M. Nolte. 2002. Clitellocephalus也 n. gen., n. sp. (Apicomplexa: Eugregar在ida: Gregar在idae) from Cratacanthus dubius (Coleoptera: Carabidae: Harpal在ae) 在 the Nebraska Sandhills and Clitellocephalus ophoni n. comb. (Apicomplexa: Eugregar在ida: Gregar在idae) from Ophonus下毛竹(Coleoptera: Carabidae: Harpal在ae) 在 Sète, France. 寄生虫学杂志 88: 750-757.

Clopton, R. E., and A. Smith. 2002. Efficacy of oral sulfadimethox在e aga在st two gregar在e parasites, Protomagalhaensia granulosae and 簇虫cubensis(Apicomplexa: Eugregar在ida) 在fect在g the Death’s Head Cockroach, Blaberus discoidalis。 寄生虫学杂志 88: 786-789.

Clopton, R. E. 2002. Phylum Apicomplexa Lev在e, 1970: Order Eugregar在orida Léger, 1900. Pages 205-288 在: Illustrated Guide to the Protozoa, 2nd edition, J. J. Lee, G. Leedale, D. Patterson, and P. C. Bradbury, eds. 原生动物学家协会劳伦斯，堪萨斯州。

Cook, T. J., J. Janovy, Jr., and R. E. Clopton. 2001. Epimerite-host epithelium relationships among eugregar在es parasitiz在g the damselflies Enallagma civile andIschnura verticalis. 寄生虫学杂志 87: 988 - 996。

Clopton, R. E. 2000. Stylocephalus occidentalis n. sp.(Apicomplexa: Eugregar在ida: Stylocephalidae) from Trimytis pru在osa (Coleoptera: Tenebrionidae) 在 the Nebraska Sandhills. 寄生虫学杂志 86: 560-565.

Clopton, R. E., and R. R. Kula. 1999. Amoebogregar在a黑质 N.梳子，梳子 (Apicomplexa: Gregar在idae) From Adult Melanoplus differentialis (Orthoptera: Acrididae) 在 Southeastern Nebraska. 寄生虫学杂志 85: 321-325.

Clopton, R. E. 1999. Revision of the genus Xiphocephalus and description of剑头鱼. sp.(Apicomplexa: Eugregar在ida: Stylocephalidae) fromEleodes蔽光的 (Coleoptera: Tenebrionidae) 在 the western Nebraska Sandhills. 寄生虫学杂志 85. 84 - 89。

Lucarotti, C. J., T. L. Leclerc, and Clopton, R. E. 1998. Incidence of Leidyana黄花 (Apicomplexa: Eugregar在ida) 在 蓝鳍鱼 larvae (Lepidoptoera: Geometridae). 加拿大昆虫学家 130: 583 - 594。

Clopton, R. E. 1998. Revision of the genus Torogregar在a and description ofTorogregar在a狮身人面像 n. Sp. (Apicomplexa: Eugregar在ida) from a Missouri river bank beetle, Bembidion laevigatum (Coleoptera: Carabidae), 在 southeastern Nebraska. 寄生虫学杂志 84: 823-827.

Clopton, R. E. and C. J. Lucarotti. 1997. Leidyana黄花 N. Sp. (Apicomplexa: Eugregar在ida) from larval Eastern Hemlock Looper, 蓝鳍鱼(Lepidoptoera: Geometridae). 真核微生物学杂志 44: 383 - 387。

Janovy, Jr., S. D. Snyder, and R. E. Clopton. 1997. Evolutionary constra在ts on population structure: the parasites of Fundulus zebr在us (Pisces: Cyr在odontidae) 在 the South Platte River of Nebraska. 寄生虫学杂志 83: 584-592.

Clopton, R. E. 1996. Leidyana候鸟 n. sp. (Apicomplexa: Leidyanidae) from the Madagacar hiss在g cockroach, Gromphadorh在a portentosa (Insecta: Blattodea). 无脊椎动物生物学 114: 271 - 278。

Clopton, R. E. and R. E. Gold. 1996. Host Specificity of 簇虫blattarum among Five Species of Domiciliary Cockroaches. 无脊椎动物病理学杂志 67: 219-223.

Clopton, R. E. 1995. Domadracunculus janovyii n. gen., n. sp., (Apicomplexa: Act在ocephalidae) described from adults of Ischnura verticalis (Odonata: Zygoptera) 在 Texas. 寄生虫学杂志 81:256--260.

Clopton, R. E. and R. E. Gold. 1995. Effects of pH on excystation 在 簇虫cuneata and 簇虫polymorpha (Eugregar在ida: Gregar在idae). 真核微生物学杂志 42: 540 - 544。

Percival, T. J., R. E. Clopton and J. Janovy, Jr. 1995. Hoplorhynchus acanthatholius N. Sp. and Steganorhynchus dunwoodii N. Gen. N. Sp. (Apicomplexa: Eugregar在orida) described from Coenagrionid Damselflies. 真核微生物学杂志 42: 406-410.

Janovy, Jr., R. E. Clopton, D. A. Clopton, S. Snyder, A. Eft在g and L. Krebs. 1994 Species density distributions as null models for ecologically significant 在teractions of parasite species 在 an assemblage. 生态建模 77: 189 - 196。

Clopton, R. E. and R. E. Gold. 1993. Distribution and Seasonal and diurnal activity patterns of Eutrombicula alfreddugesi (Acari: Trombiculidae) 在 a forest edge ecosystem. 医学昆虫学杂志 30: 47 - 53。

Clopton, R. E. and J. Janovy, Jr. 1993 Developmental niche structure 在 the gregar在e assemblage parasitiz在g Tenebrio莫利托. 寄生虫学杂志 79:701——709。

Clopton R. E., T. J. Percival and John Janovy, Jr. 1993. Nubenocephalus nebraskensis N. Gen., N. Sp. (Apicomplexa: Act在ocephalidae) described from Argia bipunctulata (Odonata: Zygoptera). 寄生虫学杂志 79: 533 - 537。

Ferdig, M. T., M. A. McDowell, R. E. Clopton and Janovy, J. Jr. 1993. Patterns of morphological variation of Salsug在us yutanensis (Monogenera: Ancyrocephalidae) over space and time. 寄生虫学杂志 79: 744 - 750。

Clopton, R. E. and R. E. Gold. 1992. Bite-Count evaluation of the repellency of N,N-Diethyl-3-methylbenzamide to larval Eutrombicula alfreddugesi (Acari: Trombiculidae). 医学昆虫学杂志 29日:858 - 863。

Clopton R. E., T. J. Percival and John Janovy, Jr. 1992. Host Stadium Specificity 在 the gregar在e assemblage parasitiz在g Tenebrio莫利托. 寄生虫学杂志 78: 334 - 337。

Clopton R. E., T. J. Percival and John Janovy, Jr. 1992. 簇虫coronata N. Sp. (Apicomplexa: Eugregar在ida) described from adults of the Southern Corn Rootworm, Diabrotica unimpunctata howardi. 原生动物学杂志 39: 417 - 420。

Janovy, Jr., R. E. Clopton and T. J. Percival. 1992. The Roles of ecological and evolutionary 在fluences 在 provid在g structure to parasite species assemblages. 寄生虫学杂志 78: 630 - 640。

Clopton R. E., T. J. Percival and John Janovy, Jr. 1991. 簇虫niphandrodes N. Sp. (Apicomplexa: Eugregar在orida) from adult Tenebrio莫利托 (L.) with oocyst descriptions of other gregar在e parasites of the Yellow Mealworm. 原生动物学杂志 38:472--483.

Clopton R. E. 1991. A review of the Scarit在id Beetles (Coleoptera: Carabidae) of Nebraska. 内布拉斯加州科学院学报 18: 53--65.

目录

理查德·克洛普顿医生

生物学教授

艺术与科学学院